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‘Neural remodelling’ in the epileptic dentate gyrus may underlie hypersynchrony and epileptogenesis.
A diffuse hyperexcitability and enhanced synchrony in the dentate gyrus (DG) are the defining characteristics of mesial temporal lobe epilepsy (MTLE). Cellular and molecular mechanisms leading to the generation of hypersynchrony have not been elucidated. A hyperactivity of principal cells in the pyramidal layer of the hippocampal DG has been suggested to be an essential mechanism. In the present study we examined if a seizure-induced elevation in extracellular glutamate (Glu) levels might influence the firing properties of DG granule cells, and, subsequently, induce a hypersynchrony in the epileptic DG. For this purpose, Glu release was elevated in the DG during the perforant path stimulation. The increase of Glu levels was induced by the injection of ibotenic acid, which was injected focally within the molecular layer of the hippocampal DG. In the present study we found a spontaneous hypersynchrony at the level of the granule cells and the hilar area in the epileptic DG. Extracellular Glu levels in the DGL were significantly increased in the epileptic DG. We investigated the effect of Glu elevation on the intrinsic firing properties of DG granule cells of controls and epileptic animals. In both control and epileptic rats, low-threshold G
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